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Anglies

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Seit ihrer wohlverdienten Rundumerneuerung präsentiert sich die Grande Dame der Hamburger Nachtclubs in prachtvollem Glanz: in edlem Dunkelblau und Gold, mit gleich zwei Tresen, an denen die gut gelaunte Barcrew die Cocktailshaker wirbelt, und — dank geöffneter Fensterfront — mit freiem Blick auf die nächtliche Reeperbahn.

You are using an outdated browser. Upgrade your browser today to experience this site better. This promoted the development of extensive lowland swamps and forests in North America and Europe.

Based on a genetic analysis of mushroom fungi, it was proposed that large quantities of wood were buried during this period because animals and decomposing bacteria and fungi had not yet evolved enzymes that could effectively digest the resistant phenolic lignin polymers and waxy suberin polymers.

They suggest that fungi that could break those substances down effectively only became dominant towards the end of the period, making subsequent coal formation much rarer.

The Carboniferous trees made extensive use of lignin. They had bark to wood ratios of 8 to 1, and even as high as 20 to 1. This compares to modern values less than 1 to 4.

Lignin is insoluble, too large to pass through cell walls, too heterogeneous for specific enzymes, and toxic, so that few organisms other than Basidiomycetes fungi can degrade it.

It can linger in soil for thousands of years and its toxic breakdown products inhibit decay of other substances.

It also may have promoted gigantism of insects and amphibians — creatures that have been constrained in size by respiratory systems that are limited in their physiological ability to transport and distribute oxygen at the lower atmospheric concentrations that have since been available.

In eastern North America, marine beds are more common in the older part of the period than the later part and are almost entirely absent by the late Carboniferous.

More diverse geology existed elsewhere, of course. Marine life is especially rich in crinoids and other echinoderms.

Brachiopods were abundant. Trilobites became quite uncommon. On land, large and diverse plant populations existed. Land vertebrates included large amphibians.

Early Carboniferous land plants, some of which were preserved in coal balls , were very similar to those of the preceding Late Devonian , but new groups also appeared at this time.

The main Early Carboniferous plants were the Equisetales horse-tails , Sphenophyllales scrambling plants , Lycopodiales club mosses , Lepidodendrales scale trees , Filicales ferns , Medullosales informally included in the " seed ferns ", an artificial assemblage of a number of early gymnosperm groups and the Cordaitales.

These continued to dominate throughout the period, but during late Carboniferous , several other groups, Cycadophyta cycads , the Callistophytales another group of "seed ferns" , and the Voltziales related to and sometimes included under the conifers , appeared.

The Carboniferous lycophytes of the order Lepidodendrales, which are cousins but not ancestors of the tiny club-moss of today, were huge trees with trunks 30 meters high and up to 1.

These included Lepidodendron with its cone called Lepidostrobus , Anabathra , Lepidophloios and Sigillaria.

The roots of several of these forms are known as Stigmaria. Unlike present-day trees, their secondary growth took place in the cortex , which also provided stability, instead of the xylem.

The fronds of some Carboniferous ferns are almost identical with those of living species. Probably many species were epiphytic.

Fossil ferns and "seed ferns" include Pecopteris , Cyclopteris , Neuropteris , Alethopteris , and Sphenopteris ; Megaphyton and Caulopteris were tree ferns.

Sphenophyllum was a slender climbing plant with whorls of leaves, which was probably related both to the calamites and the lycopods.

These plants were thought to live in swamps. True coniferous trees Walchia , of the order Voltziales appear later in the Carboniferous, and preferred higher drier ground.

In the oceans the marine invertebrate groups are the Foraminifera , corals , Bryozoa , Ostracoda , brachiopods , ammonoids , hederelloids , microconchids and echinoderms especially crinoids.

For the first time foraminifera take a prominent part in the marine faunas. The large spindle-shaped genus Fusulina and its relatives were abundant in what is now Russia, China, Japan, North America; other important genera include Valvulina , Endothyra , Archaediscus , and Saccammina the latter common in Britain and Belgium.

Some Carboniferous genera are still extant. The microscopic shells of radiolarians are found in cherts of this age in the Culm of Devon and Cornwall , and in Russia, Germany and elsewhere.

Sponges are known from spicules and anchor ropes, and include various forms such as the Calcispongea Cotyliscus and Girtycoelia , the demosponge Chaetetes , and the genus of unusual colonial glass sponges Titusvillia.

Both reef -building and solitary corals diversify and flourish; these include both rugose for example, Caninia , Corwenia , Neozaphrentis , heterocorals, and tabulate for example, Chladochonus , Michelinia forms.

Conularids were well represented by Conularia. Bryozoa are abundant in some regions; the fenestellids including Fenestella , Polypora , and Archimedes , so named because it is in the shape of an Archimedean screw.

Brachiopods are also abundant; they include productids , some of which for example, Gigantoproductus reached very large for brachiopods size and had very thick shells, while others like Chonetes were more conservative in form.

Athyridids , spiriferids , rhynchonellids , and terebratulids are also very common. Inarticulate forms include Discina and Crania.

Some species and genera had a very wide distribution with only minor variations. Annelids such as Serpulites are common fossils in some horizons.

Among the mollusca, the bivalves continue to increase in numbers and importance. Gastropods are also numerous, including the genera Murchisonia , Euomphalus , Naticopsis.

Nautiloid cephalopods are represented by tightly coiled nautilids , with straight-shelled and curved-shelled forms becoming increasingly rare.

Goniatite ammonoids are common. Trilobites are rarer than in previous periods, on a steady trend towards extinction, represented only by the proetid group.

Ostracoda , a class of crustaceans , were abundant as representatives of the meiobenthos ; genera included Amphissites , Bairdia , Beyrichiopsis , Cavellina , Coryellina , Cribroconcha , Hollinella , Kirkbya , Knoxiella , and Libumella.

Amongst the echinoderms , the crinoids were the most numerous. Dense submarine thickets of long-stemmed crinoids appear to have flourished in shallow seas, and their remains were consolidated into thick beds of rock.

Prominent genera include Cyathocrinus , Woodocrinus , and Actinocrinus. Echinoids such as Archaeocidaris and Palaeechinus were also present.

The blastoids , which included the Pentreinitidae and Codasteridae and superficially resembled crinoids in the possession of long stalks attached to the seabed, attain their maximum development at this time.

Syringothyris sp. Palaeophycus ichnosp. Crinoid calyx from the Lower Carboniferous of Ohio with a conical platyceratid gastropod Palaeocapulus acutirostre attached.

Freshwater Carboniferous invertebrates include various bivalve molluscs that lived in brackish or fresh water, such as Anthraconaia , Naiadites , and Carbonicola ; diverse crustaceans such as Candona , Carbonita , Darwinula , Estheria , Acanthocaris , Dithyrocaris , and Anthrapalaemon.

The eurypterids were also diverse, and are represented by such genera as Adelophthalmus , Megarachne originally misinterpreted as a giant spider, hence its name and the specialised very large Hibbertopterus.

Many of these were amphibious. Frequently a temporary return of marine conditions resulted in marine or brackish water genera such as Lingula , Orbiculoidea , and Productus being found in the thin beds known as marine bands.

Fossil remains of air-breathing insects , [19] myriapods and arachnids [20] are known from the late Carboniferous, but so far not from the early Carboniferous.

Their diversity when they do appear, however, shows that these arthropods were both well developed and numerous. Their large size can be attributed to the moistness of the environment mostly swampy fern forests and the fact that the oxygen concentration in the Earth's atmosphere in the Carboniferous was much higher than today.

Among the insect groups are the huge predatory Protodonata griffinflies , among which was Meganeura , a giant dragonfly -like insect and with a wingspan of ca.

Further groups are the Syntonopterodea relatives of present-day mayflies , the abundant and often large sap-sucking Palaeodictyopteroidea , the diverse herbivorous Protorthoptera , and numerous basal Dictyoptera ancestors of cockroaches.

In the Nova Scotian tree trunks land snails Archaeozonites , Dendropupa have been found. Many fish inhabited the Carboniferous seas; predominantly Elasmobranchs sharks and their relatives.

These included some, like Psammodus , with crushing pavement-like teeth adapted for grinding the shells of brachiopods, crustaceans, and other marine organisms.

Other sharks had piercing teeth, such as the Symmoriida ; some, the petalodonts , had peculiar cycloid cutting teeth. Most of the sharks were marine, but the Xenacanthida invaded fresh waters of the coal swamps.

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